# Fagus moesiaca crown foliage dry biomass, DBH (Table III lnMFC)

Foliage biomass equation for Fagus moesiaca, Greece, from Zianis & Mencuccini 2003. 16 trees, DBH 5.19–40.6 cm. Listed on Tree Allometrics, a free library of tree allometric equations, each quoted from the source that published it.

## Equation

As printed in Zianis & Mencuccini 2003 (Table III, model in caption and lnMFC row, p. 442):

> lnMFC = ln a + b lnD; ln a = −5.5168, b = 1.9979

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#equation

In this library's units, ready to compute (foliage biomass in kg):

```
Foliage biomass = 1.15 * exp(-5.5168 + 1.9979 * ln(D))
```

Inputs: D = Diameter at breast height (1.3 m) (cm).
The source's own form: 1.15 * exp(-5.5168 + 1.9979 * ln(D)), with D in cm; output in kg.
Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#canonical

## What it covers

- Output: Foliage biomass, kg
- Taxon: Fagus moesiaca (species); broadleaf
- Where: Vermio Mountain, Northern Greece. Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#where
- Biomes: temperate
- Stand: natural
- Trees measured: 16. Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#n_trees
- DBH range: 5.19–40.6 cm. Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#dbh
- R²: 0.87. Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#r2
- Residual SE: 0.5287. Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#rse
- Correction factor: 1.15 (applied). Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#cf
- Method: natural-log regression. Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#method
- Verification: read in the original source
- Last checked: 2026-11-05. Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#checked_at

A missing fit statistic is one the source doesn't state, or that we haven't recorded; it is never estimated. Outside the stated DBH range or region, the equation is an extrapolation.

## Source

Zianis, D., & Mencuccini, M. (2003). Aboveground biomass relationships for beech (Fagus moesiaca Cz.) trees in Vermio Mountain, Northern Greece, and generalised equations for Fagus sp. Annals of Forest Science, 60(5), 439–448.

- DOI: https://doi.org/10.1051/forest:2003036
- Peer review: A paper in a peer-reviewed journal.
- Access: open access
- Copy we read: https://hal.science/hal-00883716/document
- Every equation from this source: https://treeallometry.org/sources/zianis-2003/
- Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#source

## The facts, in the source's words

**Equation** (Table III, model in caption and lnMFC row, p. 442):

> lnMFC = ln a + b lnD; ln a = −5.5168, b = 1.9979

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#equation

**Equation** (Table III, lnMFC, p. 442):

> lnMFC      lnD      –5.5168      1.9979

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-expression

**Output** (Table III note, p. 442):

> MFC: foliage mass in crown

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-output

**Output** (Section 2.2, p. 441):

> oven dried to constant weight

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-output-2

**Units** (Table III note, p. 442):

> Biomass is expressed in kg, diameter in cm, and height in m.

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-published-units

**Taxon** (Abstract, p. 439):

> Fagus moesiaca Cz.

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-taxon

**Where** (Abstract, p. 439):

> Vermio Mountains of Northern
Greece

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-region

**Where** (Section 2.1, p. 440):

> the forest can be classified as temperate Mediterranean

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-region-2

**stand** (Section 2.1, p. 440 (rendered page)):

> Naturally regenerated, pure beech

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-stand

**Trees measured** (Table III caption, p. 442):

> number of sampled trees was 16

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-fit-n-trees

**Smallest DBH** (Section 2.2, p. 441):

> 5.19 to 40.6 cm

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-fit-dbh-min-cm

**Largest DBH** (Section 2.2, p. 441):

> 5.19 to 40.6 cm

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-fit-dbh-max-cm

**R²** (Table III, lnMFC, R2, p. 442):

> 0.87

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-fit-r2

**Residual standard error** (Table III, lnMFC, SEE, p. 442):

> 0.5287

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-fit-rse

**Correction factor** (Table III, lnMFC, CF, p. 442):

> 1.15

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-fit-cf

**method** (Section 3.1, p. 442):

> Napierian logarithmic function

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#quote-fit-method

**Our note:** D is DBH at 1.3 m. Multiply by Table III's CF once; it equals exp(SEE²/2), although the text calls SEE²/2 CF. Crown foliage only, not total foliage. The authors report better predictions with measured height, Eq. 5. The retained 5.19–40.6 cm range is explicitly reported in the methods (p. 441); the later comparison describes a largest Greek tree of 41.45 cm (p. 446). That sample-description discrepancy is not used to extend the reported range.

## Checks this equation passes

On every build, the library's form and the source's own form (in its units) must both give these values within 1%.

| Inputs | Expected | We get | From |
| --- | --- | --- | --- |
| D = 20 | 1.84 | 1.84 | Computed from the printed form: Computed from Table III coefficients and CF. |

Cite: https://treeallometry.org/equations/zianis-2003-iii-mfc/#checks

The calculator on the page shows carbon as biomass × 0.47 (IPCC 2006 Guidelines, Vol. 4, Ch. 4, Table 4.3) and CO₂e as carbon × 44/12.

## How to cite

Cite the source for the equation: Zianis, D., & Mencuccini, M. (2003). Aboveground biomass relationships for beech (Fagus moesiaca Cz.) trees in Vermio Mountain, Northern Greece, and generalised equations for Fagus sp. Annals of Forest Science, 60(5), 439–448. https://doi.org/10.1051/forest:2003036

Cite this page for the record and checks: Tree Allometrics (Restoration Scope), "Fagus moesiaca crown foliage dry biomass, DBH (Table III lnMFC)", checked 2026-11-05, https://treeallometry.org/equations/zianis-2003-iii-mfc/. Data CC BY 4.0 (https://creativecommons.org/licenses/by/4.0/). Quotes are the authors' words.

Data for this equation: https://treeallometry.org/data/equations/zianis-2003-iii-mfc.json
Report a problem: email info@restorationscope.com with the equation id (zianis-2003-iii-mfc) and where the right value appears.